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An interactive molecular model illustrates rotation of phi and psi angles in polypeptides, showing clashes forming and receding in real time. The Ramachandran principle is explained: The common secondary structures of alpha helices, beta strands, and turns/coil are the only conformations that are physically possible, because they avoid inter-atomic clashes. Also illustrated: the partial double-bond nature of peptide bonds and how each peptide bond holds six atoms in a plane; and the Ramachandran Plot. A short practice quiz is provided. Also available at Proteopedia.Org, and as a slideshow in English (tinyurl.com/RamachandranPrinciple) or Spanish (tinyurl.com/PrincipioRamachandran) and a YouTube video (tinyurl.com/RamachandranPrincipleYoutube). The slideshow and the video include more detailed illustrations of dihedral angles and the relation between clashes and the plot.

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